You're going to have to create a new view, or modify the current view, to accomplish this. It is a little complicated. It numbers the tracks based on the order they are listed in the view's file list.
1. Add these rules to Set rules for file display under Customize View... Use the Import/Export button to paste them in:
[Filename (path)]=[M:\Testing" [=save(0,v_ntracks_AlbumKey())1]=1 [=save(math(1+load(v_ntracks_AlbumKey())), v_ntracks_AlbumKey())1]=1 [=save(math(max(load(v_maxtn_AlbumKey()),[track #])),v_maxtn_AlbumKey())1]=1
2. Select all the tracks in the view and in the Tag Action Window, enter:
=Counter()
into the Track # field. This will assign a monotonically increasing number to each track. We'll fix this in the next step.
3. Now, with the same tracks selected, enter the following into Track # in the TAW:
=math([track #] - (load(v_maxtn_AlbumKey()) - load(v_ntracks_AlbumKey())))
This uses some calculations performed in the Set rules for file display to fix-up the track numbers, adjusting them downwards by the per-album total track count and maximum track number.
The expressions will need to be adjusted to deal with multi-disc albums though.